They had small brains; this suggests they were solitary and did not have the complex social structures of some modern cetaceans. This feature compares directly to that of modern cetaceans. Their fossils were first discovered in North Pakistan in , located at a river not far from the shores of the former Tethys Sea. The position of the nasal opening had remained unchanged since pakicetids. The fossils of ambulocetids are always found in near-shore shallow marine deposits associated with abundant marine plant fossils and littoral mollusks. Skulls of Squalodon show evidence for the first hypothesized appearance of echolocation.
This method of hearing did not give directional hearing underwater. The fossil remains of protocetids were uncovered from coastal and lagoonal facies in South Asia; unlike previous cetacean families, their fossils uncovered from Africa and North America also include open marine forms. The fossils of ambulocetids are always found in near-shore shallow marine deposits associated with abundant marine plant fossils and littoral mollusks. Wale could be heard threatening: Green Label currently attracts over twice as much traffic as MountainDew. They lived in the Eocene, approximately 48 to 35 million years ago. Archaeocetes retained aspects of their mesonychid ancestry such as the triangular teeth which modern artiodactyls, and modern whales, have lost. Since molecular analysis identifies artiodactyls as being very closely related to cetaceans, mesonychids are probably an offshoot from Artiodactyla, and cetaceans did not derive directly from them, but possibly sharing a common ancestor. Hence, ambulocetids represent a transition phase of cetacean ancestors between fresh water and marine habitat. Filter feeding is very beneficial as it allows baleen whales to efficiently gain huge energy resources, which makes the large body size in modern varieties possible. Ultimately, the change in position of the eyes and limb bones is what led the pakicetids to become waders. Hence pakicetids were most likely aquatic waders. However, it is clear that they were adapted even further to an aquatic life-style. The mandibular foramen of basilosaurids covered the entire depth of the lower jaw as in modern cetaceans. The earliest ancestors of all hoofed mammals were probably at least partly carnivorous or scavengers, and today's artiodactyls and perissodactyls became herbivores later in their evolution. Terrestrial locomotion of Rodhocetus was very limited due to their hindlimb structure. They had small brains; this suggests they were solitary and did not have the complex social structures of some modern cetaceans. Indohyus Possible relationships between cetaceans and other ungulate groups. In later species, such as Basilosaurus, the pelvic bone, no longer attached to the vertebrae and the ilium , was reduced. This is known as nasal drift. Indohyus is identified as an artiodactyl because it has two trochlea hinges , a trait unique to artiodactyls. This is demonstrated by the recovery of their fossils from a variety of coastal marine environments, including near-shore and lagoonal deposits. They too had a fluke and, unlike basilosaurids, they probably swam similarly to modern cetaceans, by using caudal oscillations. Skulls of Squalodon show evidence for the first hypothesized appearance of echolocation. In modern toothed whales, this fat pad in the mandibular foramen extends posteriorly to the middle ear. The ungulate ancestry of these archaeocetes is still underlined by characteristics like the presence of hooves at the ends of the toes in Rodhocetus. According to a study done by Spoor et al.
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